Page:Mammalia (Beddard).djvu/167

 been syndactylous, and the authors of the account of these bones think that the fourth toe may have shared in this syndactyly. The metatarsal of the fifth digit is enormously expanded at its edge, and seems to have furnished a strong support to the creature; this is also seen in the metacarpal of the fore-limb. Probably, therefore, Diprotodon was quadrupedal in its mode of progression, with the emphasis laid upon the little finger and the little toe instead of, as in ourselves, the first toe. The hind-foot of the Diprotodon could not be more unlike that of a Kangaroo than it actually is.

Another giant among these Marsupials was the genus Thylacoleo, whose name was given to it by Sir Richard Owen on the view that it was a Marsupial Tiger. Sir W. Flower has, however, controverted this opinion, and the genus is in fact, in spite of its large size, closely allied to the Phalangers and