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 320 was not published till more than half a century after Hooker first set foot in India. It is upon such foundations that Hooker's reputation as a great constructive thinker is securely based.

The first-named of these essays will probably be estimated as the most notable of them all in the History of Science. It was completed in November 1859, barely a year after the joint communications of Darwin and Wallace to the Linnean Society, and before the Origin of Species had appeared. It was to this Essay that Darwin referred when he wrote that "Hooker has come round, and will publish his belief soon." But this publication of his belief was not merely an echo of assent to Darwin's own opinions. It was a reasoned statement, advanced upon the basis of his "own self-thought," and his own wide systematic and geographical experience. From these sources he drew for himself support for the "hypothesis that species are derivative, and mutable." He points out how the natural history of Australia seemed specially suited to test such a theory, on account of the comparative uniformity of the physical features being accompanied by a great variety in its Flora, and the peculiarity of both its Fauna and Flora, as compared with other countries. After the test had been made, on the basis of study of some 8000 species, their characters, their spread, and their relations to those of other lands, he concludes decisively in favour of mutability and a doctrine of progression.

How highly this Essay was esteemed by his contemporaries is shown by the expressions of Lyell and of Darwin. The former writes: "I have just finished the reading of your splendid Essay on the Origin of Species, as illustrated by your wide botanical experience, and think it goes far to raise the variety-making hypothesis to the rank of a theory, as accounting for the manner in which new species enter the world." Darwin wrote: "I have finished your Essay. To my judgment it is by far the grandest and most interesting essay on subjects of the nature discussed I have ever read."

But besides its historical interest in relation to the Species Question, the Essay contained what was up to its time the most scientific treatment of a large area from the point of view of the