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 Rh generations, will be necessary before the naturally related groups, and their relations to one another, are recognised in the plexus of forms of the Polypodiaceae."

Such observations as these must not be understood in any sense of disparagement of the work of this great man. They are merely intended to indicate his historical position. The Origin of Species was, it is true, published some few years before the Synopsis Filicum. But we must remember that Sir William Hooker was already an old man. Few men over 70 years of age alter their opinions, and the labourer who had grown old under the belief in the Constancy of Species could not in a few brief years be expected to change the methods of thought of a long and active life. We must take Sir William Hooker as perhaps the greatest and the last of the systematists who worked under the belief in the Constancy of Species. Because we have adopted a newer point of view, and take into consideration facts and arguments which were never his, and come to different conclusions now, is no reason for valuing one whit the less the achievements of this great botanist.

His published work was just as much fundamental as was his official work. We have seen how he provided in Kew the means of indefinite development later, by constructing the coordinating machine with its collections and its libraries. In somewhat similar sense his publications were also fundamental. He did not himself construct. There is, I believe, no great modification of system or of view which is to be associated with his name. But in the wealth of trustworthy detail, recorded both pictorially and in verbal diagnoses, he has supplied the foundation for future workers to build upon, laid surely and firmly by accurate observation, and therefore durable for all time. One remark I may make as to the effect of his work on the trend of botanical activity in this country. We have noted that anatomy was not Sir William Hooker's strong point. He and many of his contemporaries did not pursue microscopic detail, and indeed seem to have avoided it. He was, however, a dominating botanical influence of the middle Victorian period. May we not see in these facts, combined with the extraordinary success of the systematic work carried on by himself, or under