Page:History of botany (Sachs; Garnsey).djvu/457

] The great majority of botanists in the second half of the 18th century had no longer any doubt that the stamens were organs of reproduction, and they were anxious to prove the existence of similar organs in the Cryptogams; they rested in this matter on external resemblances and analogies, which they interpreted in a more or less arbitrary manner. The obvious external resemblance between the antheridia and archegonia in Mosses and the sexual organs in the Phanerogams led Schmidel and Hedwig to consider them to be stamens and ovaries, and the conjecture was correct, though the true nature of the moss-fruit had to be learnt in another way. Micheli, Linnaeus and Dillen, trusting still more to external appearance and with slight knowledge of these plants, had before this taken the fruit for a male flower, and in the case of the rest of the Cryptogams the best botanists were only feeling their way in the dark with no certain experience to guide them. It is not necessary to give a particular account of the views which originated in this way; one or two may be mentioned by way of example. Koelreuter regarded the volva of Mushrooms, Gleditsch and Hedwig certain tube-like cells in their lamellae, as the male organs of fertilisation. Gleichen took the stomata, Koelreuter the indtisium, Hedwig even the glandular hairs of Ferns for anthers. It was not yet suspected that the course of development and the whole morphology of the Cryptogams could not be so compared with that of the Phanerogams; correct and incorrect assumptions with regard to the sexual organs of the Cryptogams were alike devoid of scientific value, being mere guesses and vague conjectures. Nor was the state of things much better even in the first years of the 19th century; and if by that time a number of occasional observations had been made which could afterwards be turned to scientific account, these were as yet only isolated facts without scientific connection, and every one was at liberty to concede or to refuse sexual organs to the Cryptogams generally at his own discretion. Meanwhile observations gradually accumu-