Page:History of botany (Sachs; Garnsey).djvu/286

 but he did not give a decided answer to the question. He declared however distinctly against Hedwig's reconducting vessels in the epidermis, as Sprengel had done, and it is worthy of recognition that he understood the true nature of the corners where three longitudinal walls of the parenchyma meet, while later observers found difficulties in them.

Before the appearance of Bernhardi's work the Royal scientific Society of Göttingen proposed a subject for a prize in the year 1804, which shows very plainly what uncertainty was felt at that time on all points of phytotomy. For this reason it will be well to give it at length from the preface to Rudolphi's 'Anatomic der Pflanzen' (1807): 'Since some modern physiologists deny the peculiar construction of vessels in plants which is attributed to them by other and especially the older observers, it would be well to institute new microscopical investigations, which shall either confirm the observations of Malpighi, Grew, Du Hamel, Mustel, and Hedwig, or prove that plants have a special organisation of their own which is more simple than that of animals, whether that organisation is supposed to originate in simple peculiar fibres and threads (Medicus) or with cellular and tubular tissue (tissu tubulaire of Mirbel). Attention should also be given to the following subordinate questions: 1. How many kinds of vessels may certainly be distinguished from the first period of their development? The existence of certain forms having been established; 2. Are the twisted fibres which are called spiral vessels (vasa spiralia) themselves hollow, and do they therefore form vessels, or do they serve by their convolutions for the formation of closed cavities, and how? 3. Do fluids as well as gases move in these cavities? 4. Do the scalariform ducts arise from adherence of the twisted threads (Sprengel), or do the threads owe their origin to the ducts (Mirbel)? Do alburnum and woody fibres originate in the scalariform ducts, or in true vessels, or in tubular tissue?'

We see in this case as in many similar ones, that the subject