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COLOUR SENSATION] 5. Theories of Colour-Perception.—A theory widely accepted by physicists was first proposed by Thomas Young and afterwards revived by Helmholtz. It is based on the assumption that three kinds of nervous elements exist in the retina, the excitation of which give respectively sensations of red, green and violet. These may be regarded as fundamental sensations. Homogeneous light excites all three, but with different intensities according to the length of the wave. Thus long waves will excite most strongly fibres sensitive to red, medium waves those sensitive to green, and short waves those sensitive to violet. Fig. 22 shows graphically the irritability of the three sets of fibres. Helmholtz thus applies the theory.:—

“1. Red excites strongly the fibres sensitive to red and feebly the other two—sensation: Red.

2. Yellow excites moderately the fibres sensitive to red and green, feebly the violet—sensation: Yellow.

3. Green excites strongly the green, feebly the other two—sensation: Green.

4. Blue excites moderately the fibres sensitive to green and violet, and feebly the red—sensation: Blue.

5. Violet excites strongly the fibres sensitive to violet, and feebly the other two—sensation: Violet.

6. When the excitation is nearly equal for the three kinds of fibres, then the sensation is White.”

22.—Diagram showing the Irritability of the Three Kinds of Retinal Elements.

1, red; 2, green; 3, violet. R, O, Y, G, B, V, initial letters of colours.

The Young-Helmholtz theory explains the appearance of the consecutive coloured images. Suppose, for example, that we look at a red object for a considerable time; the retinal elements sensitive to red become fatigued. Then (1) if the eye be kept in darkness, the fibres affected by red being fatigued do not act so as to give a sensation of red; those of green and of violet have been less excited, and this excitation is sufficient to give the sensation of pale greenish blue; (2) if the eye be fixed on a white surface, the red fibres, being fatigued, are not excited by the red rays contained in the white light; on the contrary, the green and violet fibres are strongly excited, and the consequence is that we have an intense complementary image; (3) if we look at a bluish green surface, the complementary of red, the effect will be to excite still more strongly the green and violet fibres, and consequently to have a still more intense complementary image; (4) if we regard a red surface, the primitive colour, the red fibres are little affected in consequence of being fatigued, the green and violet fibres will be only feebly excited, and therefore only a very feeble complementary image will be seen; and (5) if we look at a surface of a different colour altogether, this colour may combine with that of the consecutive image, and produce a mixed colour, thus, on a yellow surface, we will see an image of an orange colour.

Every colour has three qualities: (1) hue, or tint, such as red, green, violet; (2) degree of saturation, or purity, according to the amount of white mixed with the tint, as when we recognize a red or green as pale or deep; and (3) intensity, or luminosity, or brightness as when we designate the tint of a red rose as dark or bright. Two colours are identical when they agree as to these three qualities. Observation shows, however, that out of one hundred men ninety-six agree in identifying or in discriminating colours, while the remaining four show defective appreciation. These latter are called colour-blind. This defect is about ten times less frequent in women. Colour-blindness is congenital and incurable, and it is due to an unknown condition of the retina or nerve centres, or both, and must be distinguished from transient colour-blindness, sometimes caused by the excessive use of tobacco and by disease. When caused by tobacco, the sensation of blue is the last to disappear. Absolute inability to distinguish colour is rare, if it really exists; in some rare cases there is only one colour sensation; and in a few

cases the colour-blind fails to distinguish blue from green, or there is insensibility to violet. Daltonism, or red-green blindness, of which there are two varieties, the red-blind and the green-blind, is the more common defect. Red appears to a redblind person as a dark green or greenish yellow, yellow and orange as dirty green, and green is green and brighter than the green of the yellow and orange. To a green-blind person red appears as dark yellow, yellow is yellow, except a little lighter in shade than the red he calls dark yellow, and green is pale yellow.

According to the Young-Helmholtz theory, there are three fundamental colour sensations, red, green and violet, by the combination of which all other colours may be formed, and it is assumed that there exist in the retina three kinds of nerve elements, each of which is specially responsive to the stimulus of waves of a certain frequency corresponding to one colour, and much less so to waves of other frequencies and other colours. If waves corresponding to pure red alone act on the retina, only the corresponding nerve element for red would be excited, and so with green and violet. But if waves of different frequencies are mixed (corresponding to a mixture of colours), then the nerve elements will be set in action in proportion to the amount and intensity of the constituent excitant rays in the colour. Thus if all the nerve elements were simultaneously set in action, the sensation is that of white light; if that corresponding to red and green, the resultant sensation will be orange or yellow; if mainly the green and violet, the sensation will be blue and indigo. Then red-blindness may be explained by supposing that the elements corresponding to the sensation of red are absent; and green-blindness, to the absence of the elements sensitive to green. If to a red-blind person the green and violet are equal, and when to a green-blind person the red and violet are equal, they may have sensations which to them constitute white, while to the normal eye the sensation is not white, but bluish green in the one case and green in the other. In each case, to the normal eye, the sensation of green has been added to the sensations of red and blue. It will be evident, also, that whiteness to the colour-blind eye cannot be the same as whiteness to the normal eye. No doubt this theory explains certain phenomena of colour-blindness, of after-coloured images, and of contrast of colour, but it is open to various objections. It has no anatomical basis, as it has been found to be impossible to demonstrate the existence of three kinds of nerve elements, or retinal elements, corresponding to the three fundamental colour sensations. Why should red to a colour-blind person give rise to a sensation of something like green, or why should it give rise to a sensation at all? Again, and as already stated, in cases of colour-blindness due to tobacco or to disease, only blue may be seen, while it is said that the rest of the spectrum seems to be white. It is difficult to understand how white can be the sensation if the sensations of red and green are lost. On the other hand, it may be argued that such colour-blind eyes do not really see white as seen by a normal person, and that they only have a sensation which they have been accustomed to call white. According to this theory, we never actually experience the primary sensations. Thus we never see primary red, as the sensation is more or less mixed with primary green, and even with primary blue (violet). So with regard to primary green and primary violet. Helmholtz, in his last work on the subject, adopted as the three primary colours a red bluer than spectral red, (a) a green lying between 540 λ and 560 λ (b, like the green of vegetation), and a blue at about 470 λ (c, like ultramarine), all, however, much more highly saturated than any colours existing in the spectrum.

In Handbuch der Physiologischen Optik (Hamburg and Leipzig, 1896) Helmholtz pointed out that luminosity or brightness plays a more important part in colour perception than has been supposed. Each spectral colour is composed of certain proportions of these fundamental colours, or, to put it in another way, a combination of two of them added to a certain amount of white.

Hering's theory proceeds on the assumption of chemical changes in the retina under the influence of light. It also assumes that certain fundamental sensations are excited by light or occur during the absence of light. These fundamental sensations are white, black, red, yellow, green and blue. They are arranged in pairs, the one colour in each pair being, in a sense, complementary to the other, as white to black, red to green, and yellow to blue. Hering also supposes that when rays of a certain wave-length fall on visual substances assumed to exist in the retina, destructive or, as it is termed, katabolic changes occur, while rays having other wave-lengths cause constructive or anabolic changes. Suppose that in a red-green substance katabolic and anabolic changes occur in equal amount, there may be no sensation, but when waves of a certain wave-length or frequency cause katabolic changes in excess, there will be a sensation of red, while shorter waves and of greater frequency, by exciting anabolic changes, will cause a sensation of green. In like manner, katabolism of a yellow-blue visual substance gives rise to a sensation we call yellow, while anabolism by shorter waves acting on the same substance, causes the sensation of blue. Again, katabolism of a white-black visual substance