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PALAEONTOLOGICAL] in question, which is called by the name of the most abundant species. In this way what is called a “geological horizon,” or “zone,” is marked off, and its exact position in the series of formations is fixed.

Perhaps the most distinctive feature in the progress of palaeontological geology during the last half century has been the recognition and wide application of this method of zonal stratigraphy, which, in itself, was only a further development of William Smith’s famous idea, “Strata identified by Organized Fossils.” It was first carried out in detail by various palaeontologists in reference to the Jurassic formations, notably by F. A. von Quenstedt and C. A. Oppel in Germany and d’Orbigny in France. The publication of Oppel’s classic work Die Juraformation Englands, Frankreichs und des südwestlichen Deutschlands (1856–1858) marked an epoch in the development of stratigraphical geology. Combining what had been done by various observers with his own laborious researches in France, England, Württemberg and Bavaria, he drew up a classification of the Jurassic system, grouping its several formations into zones, each characterized by some distinctly predominant fossil after which it was named (see ). The same method of classification was afterwards extended to the Cretaceous series by A. D. d’Orbigny, E. Hébert and others, until the whole Mesozoic rocks from the Trias to the top of the Chalk has now been partitioned into zones, each named after some characteristic species or genus of fossils. More recently the principle has been extended to the Palaeozoic formations, though as yet less fully than to the younger parts of the geological record. It has been successfully applied by Professor C. Lapworth to the investigation of the Silurian series (see ; ). He found that the species of graptolites have each a comparatively narrow vertical range, and they may consequently be used for stratigraphical purposes. Applying the method, in the first instance, to the highly plicated Silurian rocks of the south of Scotland, he found that by means of graptolites he was able to work out the structure of the ground. Each great group of strata was seen to possess its own graptolitic zones, and by their means could be identified not only in the original complex Scottish area, but in England and Wales and in Ireland. It was eventually ascertained that the succession of zones in Great Britain could be recognized on the Continent, in North America and even in Australia. The brachiopods and trilobites have likewise been made use of for zonal purposes among the oldest sedimentary formations. The most ancient of the Palaeozoic systems has as its fitting base the Olenellus zone.

Within undefined and no doubt variable geographical limits palaeontological zones have been found to be remarkably persistent. They follow each other in the same general order, but not always with equal definiteness. The type fossil may appear in some districts on a higher or a lower platform than it does in others. Only to a limited degree is there any coincidence between lithological variations in the strata and the sequence of the zones. In the Jurassic formations, indeed, where frequent alternations of different sedimentary materials are to be met with, it is in some cases possible to trace a definite upward or downward limit for a zone by some abrupt change in the sedimentation, such as from limestone to shale. But such a precise demarcation is impossible where no distinct bands of different sediments are to be seen. The zones can then only be vaguely determined by finding their characteristic fossils, and noting where these begin to appear in the strata and where they cease. It would seem, therefore, that the sequence of palaeontological zones, or life-horizons, has not depended merely upon changes in the nature of the conditions under which the organisms lived. We should naturally expect that these changes would have had a marked influence; that, for instance, a difference should be perceptible between the character of the fossils in a limestone and that of those in a shale or a sandstone. The environment, when a limestone was in course of deposition, would generally be one of clear water, favourable for a more vigorous and more varied fauna than where a shale series was accumulating, when the water would be discoloured, and only such animals would continue to live in it, or on the bottom, as could maintain themselves in the midst of mud. But no such lithological reason, betokening geographical changes that would affect living creatures, can be adduced as a universally applicable explanation of the occurrence and limitation of palaeontological zones. One of these zones may be only a few inches, or feet or yards in vertical extent, and no obvious lithological or other cause can be seen why its specially characteristic fossils should not be found just as frequently in the similar strata above and below. There is often little or no evidence of any serious change in the conditions of sedimentation, still less of any widespread physical disturbance, such as the catastrophes by which the older geologists explained the extinction of successive types of life.

It has been suggested that, where the life-zones are well defined, sedimentation has been extremely slow, and that though these zones follow each other with no break in the sedimentation, they were really separated by prolonged intervals of time during which organic evolution could come effectively into play. But it is not easy to explain how, for example in the Lower Lias, there could have been a succession of prodigious intervals, when practically no sediment was laid down, and yet that the strata should show no sign of contemporaneous disturbance or denudation, but succeed each other as if they had been accumulated by one continuous process of deposit. It must be admitted that the problem of life-zones in stratigraphical geology has not yet been solved.

As Darwin first cogently showed, the history of life has been very imperfectly registered in the stratified parts of the earth’s crust. Apart from the fact that, even under the most favourable conditions, only a small proportion of the total flora and fauna of any period would be preserved in the fossil state, enormous gaps occur where no record has survived at all. It is as if whole chapters and books were missing from a historical work. Some of these lacunae are sufficiently obvious. Thus, in some cases, powerful dislocations have thrown considerable portions of the rocks out of sight. Sometimes extensive metamorphism has so affected them that their original characters, including their organic contents, have been destroyed. Oftenest of all, denudation has come into play, and vast masses of fossiliferous rock have been entirely worn away, as is demonstrated by the abundant unconformabilities in the structure of the earth’s crust.

While the mere fact that one series of rocks lies unconformably on another proves the lapse of a considerable interval between their respective dates, the relative length of this interval may sometimes be proved by means of fossil evidence, and by this alone. Let us suppose, for example, that a certain group of formations has been disturbed, upraised, denuded and covered unconformably by a second group. In lithological characters the two may closely resemble each other, and there may be nothing to show that the gap represented by their unconformability is of an important character. In many cases, indeed, it would be quite impossible to pronounce any well-grounded judgment as to the amount of interval, even measured by the vague relative standards of geological chronology. But if each group contains a well-preserved suite of organic remains, it may not only be possible, but easy, to say exactly how much of the geological record has been left out between the two sets of formations. By comparing the fossils with those obtained from regions where the geological record is more complete, it may be ascertained, perhaps, that the lower rocks belong to a certain platform or stage in geological history which for our present purpose we may call D, and that the upper rocks can in like manner be paralleled with stage H. It would be then apparent that at this locality the chronicles of three great geological periods E, F, and G were wanting, which are elsewhere found to be intercalated between D and H. The lapse of time represented by this unconformability would thus be equivalent to that required for the accumulation of the three missing formations in those regions where sedimentation was more continuous.

Fossil evidence may be made to prove the existence of gaps which are not otherwise apparent. As has been already remarked, changes in organic forms must, on the whole, have been extremely slow in the geological past. The whole species of a sea-floor could not pass entirely away, and be replaced by other forms, without the lapse of long periods of time. If then among the conformable stratified formations of former ages we encounter sudden and abrupt changes in the facies of the fossils, we may be certain that these must mark omissions in the record, which we may hope to fill in from a more perfect series elsewhere. The complete biological contrasts between the fossil contents of unconformable strata are sufficiently explicable. It is not so easy to give a satisfactory account of those which occur where the beds are strictly conformable, and where no evidence can be observed of any considerable change of physical conditions at the time of deposit. A group of strata having the same general lithological characters throughout may be marked by a great discrepance between the fossils above and below a certain line. A few species may pass from the one into the other, or perhaps every species may be different. In cases of this kind, when proved to be not merely local but persistent over wide areas, we must admit, notwithstanding the apparently undisturbed and continuous character of the original deposition of the strata, that the abrupt transition from the one facies of fossils to the other represents a long interval of time which has not been recorded by the deposit of strata. A. C. Ramsay, who called attention to these gaps, termed them “breaks in the succession of organic remains.” He showed that they occur abundantly among the Palaeozoic and Secondary rocks of England. It is obvious, of course, that such breaks, even though traceable over wide regions, were not general over the whole globe. There have never been any universal interruptions in the continuity of the chain of being, so far as geological evidence can show. But the physical changes which caused the breaks may have been general over a zoological district or minor region. They no doubt often caused the complete extinction of genera and species which had a small geographical range.

From all these facts it is clear that the geological record, as it now exists, is at the best but an imperfect chronicle of geological history. In no country is it complete. The lacunae of one region must be supplied from another. Yet in proportion to the geographical distance between the localities where the gaps occur and those whence the missing intervals are supplied, the element of uncertainty in our reading of the record is increased. The most desirable method of research is to exhaust the evidence for each area or province, and to compare the general order of its succession as a whole with that which can be established for other provinces.