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 FLOURENS, GUSTAVE (1838–1871), French revolutionist and writer, a son of J. P. Flourens (1794–1867), the physiologist, was born at Paris on the 4th of August 1838. In 1863 he undertook for his father a course of lectures at the Collège de France, the subject of which was the history of mankind. His theories as to the manifold origin of the human race, however, gave offence to the clergy, and he was precluded from delivering a second course. He then went to Brussels, where he published his lectures under the title of Histoire de l’homme (1863); he next visited Constantinople and Athens, took part in the Cretan insurrection of 1866, spent some time in Italy, where an article of his in the Popolo d’Italia caused his arrest and imprisonment, and finally, having returned to France, nearly lost his life in a duel with Paul de Cassagnac, editor of the Pays. In Paris he devoted his pen to the cause of republicanism, and at length, having failed in an attempt to organize a revolution at Belleville on the 7th of February 1870, found himself compelled to flee from France. Returning to Paris on the downfall of Napoleon, he soon placed himself at the head of a body of 500 tirailleurs. On account of his insurrectionary proceedings he was taken prisoner at Créteil, near Vincennes, by the provisional government, and confined at Mazas on the 7th of December 1870, but was released by his men on the night of January 21–22. On the 18th of March he joined the Communists. He was elected a member of the commune by the 20th arrondissement, and was named colonel. He was one of the most active leaders of the insurrection, and in a sortie against the Versailles troops in the morning of the 3rd of April was killed in a hand-to-hand conflict at Rueil, near Malmaison. Besides his Science de l’homme (Paris, 1869), Gustave Flourens was the author of numerous fugitive pamphlets.

See C. Prolès, Les Hommes de la révolution de 1871 (Paris, 1898).

 FLOURENS, MARIE JEAN PIERRE (1794–1867), French physiologist, was born at Maureilhan, near Béziers, in the department of Hérault, on the 15th of April 1794. At the age of fifteen he began the study of medicine at Montpellier, where in 1823 he received the degree of doctor. In the following year he repaired to Paris, provided with an introduction from A. P. de Candolle, the botanist, to Baron Cuvier, who received him kindly, and interested himself in his welfare. At Paris Flourens engaged in physiological research, occasionally contributing to literary publications; and in 1821, at the Athénée there, he gave a course of lectures on the physiological theory of the sensations, which attracted much attention amongst men of science. His paper entitled Recherches expérimentales sur les propriétés et les fonctions du système nerveux dans les animaux vertébrés, in which he, from experimental evidence, sought to assign their special functions to the cerebrum, corpora quadrigemina and cerebellum, was the subject of a highly commendatory report by Cuvier, adopted by the French Academy of Sciences in 1822. He was chosen by Cuvier in 1828 to deliver for him a course of lectures on natural history at the Collège de France, and in the same year became, in succession to L. A. G. Bosc, a member of the Institute, in the division “Économie rurale.” In 1830 he became Cuvier’s substitute as lecturer on human anatomy at the Jardin du Roi, and in 1832 was elected to the post of titular professor, which he vacated for the professorship of comparative anatomy created for him at the museum of the Jardin the same year. In 1833 Flourens, in accordance with the dying request of Cuvier, was appointed a perpetual secretary of the Academy of Sciences; and in 1838 he was returned as a deputy for the arrondissement of Béziers. In 1840 he was elected, in preference to Victor Hugo, to succeed J. F. Michaud at the French Academy; and in 1845 he was created a commander of the legion of honour, and in the next year a peer of France. In March 1847 Flourens directed the attention of the Academy of Sciences to the anaesthetic effect of chloroform on animals. On the revolution of 1848 he withdrew completely from political life; and in 1855 he accepted the professorship of natural history at the Collège de France. He died at Montgeron, near Paris, on the 6th of December 1867.

Besides numerous shorter scientific memoirs, Flourens published—Essai sur quelques points de la doctrine de la révulsion et de la dérivation (Montpellier, 1813); Expériences sur le système nerveux (Paris, 1825); Cours sur la génération, l’ovologie, et l’embryologie (1836); Analyse raisonnée des travaux de G. Cuvier (1841); Recherches sur le développement des os et des dents (1842); Anatomie générale de la peau et des membranes muqueuses (1843); Buffon, histoire de ses travaux et de ses idées (1844); Fontenelle, ou de la philosophie moderne relativement aux sciences physiques (1847); Théorie expérimentale de la formation des os (1847); Œuvres complètes de Buffon (1853); De la longévité humaine et de la quantité de vie sur le globe (1854), numerous editions; Histoire de la découverte de la circulation du sang (1854); Cours de physiologie comparée (1856); Recueil des éloges historiques (1856); De la vie et de l’intelligence (1858); De la raison, du génie, et de la folie (1861); Ontologie naturelle (1861); Examen du livre de M. Darwin sur l’Origine des Espèces (1864). For a list of his papers see the Royal Society’s Catalogue of Scientific Papers.

 FLOWER, SIR WILLIAM HENRY (1831–1899), English biologist, was born at Stratford-on-Avon on the 30th of November 1831. Choosing medicine as his profession, he began his studies at University College, London, where he showed special aptitude for physiology and comparative anatomy and took his M.B. degree in 1851. He then joined the Army Medical Service, and went out to the Crimea as assistant-surgeon, receiving the medal with four clasps. On his return to England he became a member of the surgical staff of the Middlesex hospital, London, and in 1861 succeeded J. T. Quekett as curator of the Hunterian Museum of the Royal College of Surgeons of England. In 1870 he also became Hunterian professor, and in 1884, on the death of Sir Richard Owen, was appointed to the directorship of the Natural History Museum at South Kensington. He died in London on the 1st of July 1899. He made valuable contributions to structural anthropology, publishing, for example, complete and accurate measurements of no less than 1300 human skulls, and as a comparative anatomist he ranked high, devoting himself especially to the study of the mammalia. He was also a leading authority on the arrangement of museums. The greater part of his life was spent in their administration, and in consequence he held very decided views as to the principles upon which their specimens should be set out. He insisted on the importance of distinguishing between collections intended for the use of specialists and those designed for the instruction of the general public, pointing out that it was as futile to present to the former a number of merely typical forms as to provide the latter with a long series of specimens differing only in the most minute details. His ideas, which were largely and successfully applied to the museums of which he had charge, gained wide approval, and their influence entitles him to be looked upon as a reformer who did much to improve the methods of museum arrangement and management. In addition to numerous original papers, he was the author of An Introduction to the Osteology of the Mammalia (1870); Fashion in Deformity (1881); The Horse: a Study in Natural History (1890); Introduction to the Study of Mammals, Living and Extinct (1891); Essays on Museums and other Subjects (1898). He also wrote many articles for the ninth edition of the Encyclopaedia Britannica.  FLOWER (Lat. flos, floris; Fr. fleur), a term popularly used for the bloom or blossom of a plant, and so by analogy for the fairest, choicest or finest part or aspect of anything, and in various technical senses. Here we shall deal only with its botanical interest. It is impossible to give a rigid botanical definition of the term “flower.” The flower is a characteristic feature of the highest group of the plant kingdom—the flowering plants (Phanerogams)—and is the name given to the association of organs, more or less leaf-like in form, which are concerned with the production of the fruit or seed. In modern botanical works the group is often known as the seed-plants (Spermatophyta). As the seed develops from the ovule which has been fertilized by the pollen, the essential structures for seed-production are two, viz. the pollen-bearer or stamen and the ovule-bearer or carpel. These are with few exceptions foliar structures, known in comparative morphology as sporophylls, because they bear the spores, namely, the microspores or pollen-grains which are developed in the microsporangia or pollen-sacs, and the megaspore, which is contained in the ovule or megasporangium.

In (q.v.), which represent the more primitive 