Page:EB1911 - Volume 10.djvu/525

Rh remarkable circumstance) they are always adapted to the speed at which the wing is travelling for the time being. The increase and decrease in the angles made by the wing as it hastens to and fro are due partly to the resistance offered by the air, and partly to the mechanism and mode of application of the wing to the air. The wing, during its vibrations, rotates upon two separate centres, the tip rotating round the root of the wing as an axis (short axis of wing), the posterior margin rotating around the anterior margin (long axis of wing). The wing is really eccentric in its nature, a remark which applies also to the rowing feathers of the bird’s wing. The compound rotation goes on throughout the entire down and up strokes, and is intimately associated with the power which the wing enjoys of alternately seizing and evading the air.

The compound rotation of the wing is greatly facilitated by the wing being elastic and flexible. It is this which causes the wing to twist and untwist diagonally on its long axis when it is made to vibrate. The twisting referred to is partly a vital and partly a mechanical act;—that is, it is occasioned in part by the action of the muscles and in part by the greater resistance experienced from the air by the tip and posterior margin of the wing as compared with the root and anterior margin,—the resistance experienced by the tip and posterior margin causing them to reverse always subsequently to the root and anterior margin, which has the effect of throwing the anterior and posterior margins of the wing into figure-of-8 curves, as shown at figs. 9, 11, 12, 16, 18, 20, 21, 22 and 23.

The compound rotation of the wing, as seen in the bird, is represented in fig. 24.

Not the least curious feature of the wing movements is the remarkable power which the wing possesses of making and utilizing its own currents. Thus, when the wing descends it draws after it a strong current, which, being met by the wing during its ascent, greatly increases the efficacy of the up stroke. Similarly and conversely, when the wing ascends, it creates an upward current, which, being met by the wing when it descends, powerfully contributes to the efficiency of the down stroke. This statement can be readily verified by experiment both with natural and artificial wings. Neither the up nor the down strokes are complete in themselves.

The wing to act efficiently must be driven at a certain speed, and in such a manner that the down and up strokes shall glide into each other. It is only in this way that the air can be made to pulsate, and that the rhythm of the wing and the air waves can be made to correspond. The air must be seized and let go in a certain order and at a certain speed to extract a maximum recoil. The rapidity of the wing movements is regulated by the size of the wing, small wings being driven at a very much higher speed than larger ones. The different parts of the wing, moreover, travel at different degrees of velocity—the tip and posterior margin of the wing always rushing through a much greater space, in a given time, than the root and anterior margin.

The rapidity of travel of the insect wing is in some cases enormous. The wasp, for instance, is said to ply its wings at the rate of 110, and the common house-fly at the rate of 330 beats per second. Quick as are the vibrations of natural wings, the speed of certain parts of the wing is amazingly increased. Wings as a rule are long and narrow. As a consequence, a comparatively slow and very limited movement at the root confers great range and immense speed at the tip, the speed of each portion of the wing increasing as the root of the wing is receded from. This is explained on a principle well understood in mechanics, viz. that when a wing or rod hinged at one end is made to move in a circle, the tip or free end of the wing or rod describes a much wider circle in a given time than a portion of the wing or rod nearer the hinge (fig. 25).

One naturally inquires why the high speed of wings, and why the progressive increase of speed at their tips and posterior margins? The answer is not far to seek. If the wings were not driven at a high speed, and if they were not eccentrics made to revolve upon two separate axes, they would of necessity be large cumbrous structures; but large heavy wings would be difficult to work, and what is worse, they would (if too large), instead of controlling the air, be controlled by it, and so cease to be flying organs.

There is, however, another reason why wings should be made to vibrate at high speeds. The air, as explained, is a very light, thin, elastic medium, which yields on the slightest pressure, and unless the wings attacked it with great violence the necessary recoil or resistance could not be obtained. The atmosphere, because of its great tenuity, mobility and comparative imponderability, presents little resistance to bodies passing through it at low velocities. If, however, the speed be greatly accelerated,