Page:EB1911 - Volume 08.djvu/954

 premaxillary bone and is often referred to a genus apart, as Zäedius.

In Tatoua (Cabassous or Lysiurus) the teeth are or, of moderate size and subcylindrical: the most posterior placed a little way behind the anterior root of the zygoma, but far from the hinder margin of the palate. Skull somewhat elongated, much constricted behind the orbits, and immediately in front of the constriction considerably dilated. Lower jaw slender, with the coronoid process small and sharp pointed, sometimes obsolete. Vertebrae: C 7, D 12-13, L 5, S 10, Ca 18. Head broad behind. Ears rather large and rounded, wide apart. Movable bands of armour 12-13. Tail considerably shorter than the body, and slender, covered with nearly naked skin, with a few small, scattered, bony plates, chiefly on the under surface and near the apex. On the fore-feet the first and second toes are long and slender, with small claws and the normal number of phalanges. The other toes have but two phalanges; the third has an immense sickle-like claw; the fourth and fifth similar but smaller claws. The hind-feet are comparatively small, with five toes, and small, triangular, blunt nails; the third longest, the first shortest. The best-known species of this genus, the tatouay or cabassou, T. unicinctus, is, after Priodon gigas, the largest of the group. It is found, though not abundantly, in Surinam, Brazil and Paraguay. Others, such as T. hispidus and T. lugubris, have been described.

In the giant armadillo (Priodon gigas) the teeth are variable in number, and generally differ on the two sides of each jaw, being usually from 20 to 25 on each side above and below, so that as many as a hundred may be present altogether; but as life advances the anterior teeth fall out, and all traces of their sockets disappear. The series extends as far back as the hinder edge of the anterior root of the zygoma. They are all very small, in the anterior half of each series strongly compressed, with flat sides and a straight free edge, but posteriorly more cylindrical, with flat, truncated, free surfaces. Vertebrae: C 7, D 12, L 3, S 10, Ca 23. Head small, elongated, conical. Ears moderate, ovate. Armour with 12-13 movable bands. Tail nearly equal to the body in length, gradually tapering, closely covered with quadrangular scales, arranged in a quincunx pattern. Fore-feet with five toes, formed on the same plan as those of Tatoua, but with the claw of the third of still greater size, and that of the others, especially the fifth, proportionally reduced. Hind-foot short and rounded, with five very short toes, and short, broad, flat obtuse nails. The giant armadillo is by far the largest existing member of the family, measuring rather more than 3 ft. from the tip of the nose to the root of the tail, the tail being about 20 in. long. It inhabits the forest of Surinam and Brazil. The powerful claws of its fore-feet enable it to dig with great facility; and its food consists chiefly of termites and other insects, although it is said to attack and uproot newly-made graves for the purpose of devouring the flesh of the bodies contained in them.

The apar (Tolypeutes tricinctus) typifies a genus in which the teeth are or, and are rather large in proportion to the size of the skull, with the hinder end of the series reaching nearly to the posterior margin of the palate. Vertebrae: C 7, D 11, L 3, S 12, Ca 13. Ears placed low on the sides of the head, rather large, broadly ovate. Armour with its scapular and pelvic shields very free at the sides of the body, forming large chambers into which the limbs can be readily withdrawn, and only three movable bands. Tail short, conical, covered with large bony tubercles. The fore-feet formed on the same type as in the last genus, but the peculiarities carried to a still greater extent. The claw of the third toe is very long, while those of the first and fifth are greatly reduced and sometimes wanting. On the hind-foot the three middle toes have broad, flat, subequal nails, forming together a kind of tripartite hoof; the first and fifth much shorter, with more compressed nails.

The armadillos of this genus have the power of rolling themselves up into a ball, the shield on the top of the head and the tuberculated dorsal surface of the tail exactly fitting into and filling up the apertures left by the notches at either end of the body-armour. This appears to be their usual means of defence when frightened or surprised, as they do not burrow like the other species. They run very quickly, with a very peculiar gait, only the tips of the claws of the fore-feet touching the ground. In addition to the apar, there are the Argentine and Bolivian T. conurus, and T. muriei from Argentina or Patagonia.

The last group of existing armadillos forms the genus Tatusia and the subfamily Tatusiinae; the subfamily rank being based on the fact that of the seven or eight pairs of small subcylindrical teeth, all but the last, which is considerably smaller than the rest, are preceded by milk-teeth not changed until the animal has nearly attained full size. Vertebrae: C 7, D 9-11, L 5, S 8, Ca 20-27. Head narrow, with a long, narrow, subcylindrical obliquely truncated snout. Ears rather large, ovate and erect, placed close together on the occiput. Armour with seven to nine distinct movable bands. Body generally elongated and narrow. Tail moderate, or long, gradually tapering; its plates forming distinct rings for the greater part of its length. Fore-feet with four visible toes, and a concealed clawless rudiment of the fifth; the claws long, slightly curved, and slender, the third and fourth subequal and alike, the first and fourth much shorter. Hind-feet with five toes, armed with strong, slightly curved, conical, obtusely pointed nails, and the third longest, then the second and fourth, and the first and fifth much shorter than the others. This genus differs from all the other armadillos in having a pair of inguinal teats in addition to the usual pectoral pair, and in producing a large number (4 to 10) of young at a birth, all the others having usually but one or two. The peba armadillo, T. septemcincta, is a well-known species, having an extensive range from Texas to Paraguay. It is replaced in the more southern regions of South America by a smaller species, with shorter tail, the mulita (T. hybrida) so called from the resemblance of its head and ears to those of a mule. T. kappleri is a large species from Guiana.

Finally we have the pichiciago, or fairy armadillo, Chlamydophorus truncatus, typifying the subfamily Chlamydophorinae. In most anatomical characters, especially the structure of the fore-foot, this group resembles the Dasypodinae, but it differs remarkably from all other known armadillos, living or extinct, in the peculiar modification of the armour.

The teeth, which number, are subcylindrical, somewhat compressed, moderate in size, and smaller at each end (especially in front) than at the middle of the series. Skull broad and rounded behind, pointed in front. Muzzle subcylindrical and depressed. A conspicuous rounded rough prominence on the frontal bone, just before each orbit. Tympanic prolonged into a tubular auditory meatus, curving upwards round the base of the zygoma. Vertebrae: C 7, D 11, L 3, S 10, Ca 15. Upper part of head and trunk covered with four-sided horny plates (with small thin ossifications beneath), forming a shield, free and overhanging the sides of the trunk, and attached only along the middle line of the back. The plates are arranged in a series of distinct transverse bands, about twenty in number between the occiput and the posterior truncated end, and not divided into solid scapular and pelvic shields with movable bands between. The hinder end of the body is abruptly truncated and covered by a vertically placed, strong, solid, bony shield, of an oval (transversely extended) form, covered by thin horny plates. This shield is firmly welded by five bony processes to the hinder part of the pelvis. Through a notch in the middle of its lower border the tail passes out. The latter is rather short, cylindrical in its proximal half, and expanded and depressed or spatulate in its terminal portion, and covered with horny plates. The dorsal surfaces of the fore and hind-feet are also covered with horny plates. The remainder of the limbs and under surface and sides of the body beneath the overlapping lateral parts of the back shield are clothed with rather long, soft silky hair. Eyes and ears very small, and concealed by the hair. Extremities short. Feet large, each with five well-developed claws, those on the fore-feet very long, stout and subcompressed, the structure of the digits being essentially the same as those of Tatoua and Priodon. Teats two, pectoral. Visceral anatomy closely