Page:EB1911 - Volume 08.djvu/235

 expressed in calories, one calory being the amount of heat necessary to raise the temperature of one kilogram of water one degree centigrade. But it is to be observed that this unit is employed simply from convenience, and without implication as to what extent the energy of food is converted into heat in the body. The unit employed in the measurement of some other form of energy might be used instead, as, for example, the foot-ton, which represents the amount of energy necessary to raise one ton through one foot.

The amount of energy which a given quantity of food will produce on complete oxidation outside the body, however, is greater than that which the body will actually derive from it. In the first place, as previously shown, part of the food will not be digested and absorbed. In the second place, the nitrogenous compounds absorbed are not completely oxidized in the body, the residuum being excreted in the urine as urea and other bodies that are capable of further oxidation in the calorimeter. The total heat of combustion of the food eaten must therefore be diminished by the heat of combustion of the oxidizable material rejected by the body, to find what amount of energy is actually available to the organism for the production of work and heat. The amount thus determined is commonly known as the fuel value of food.

Rubner’s commonly quoted estimates for the fuel value of the nutrients of mixed diet are,—for protein and carbohydrates 4·1, and for fats 9·3 calories per gram. According to the method of deduction, however, these factors were more applicable to digested than to total nutrients. Atwater and associates have deduced, from data much more extensive than those available to Rubner, factors for total nutrients somewhat lower than these, as shown in Table III. These estimates seem to represent the best average factors at present available, but are subject to revision as knowledge is extended.