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268 inequality in the female on the opposite sides of the body. In Gelasimus, according to a statement quoted by Milne Edwards, the male and the female live in the same burrow, and this shews that they pair; the male closes the mouth of the burrow with one of its chelæ, which is enormously developed; so that here it indirectly serves as a means of defence. Their main use, however, is probably to seize and to secure the female, and this in some instances, as with Gammarus, is known to be the case. The male of the hermit or soldier crab (Pagurus) for weeks together, carries about the shell inhabited by the female. The sexes, however, of the common shore-crab (Carcinus mænas), as Mr. Bate informs me, unite directly after the female has moulted her hard shell, when she is so soft that she would be injured if seized by the strong pincers of the male; but as she is caught and carried about by the male before moulting, she could then be seized with impunity.

Fritz Müller states that certain species of Melita are distinguished from all other amphipods by the females having "the coxal lemellæ of the penultimate pair of feet produced into hook-like processes, of which the males lay hold with the hands of the first pair." The development of these hook-like processes has probably followed from those females which were the most securely held during the act of reproduction, having left the largest number of offspring. Another Brazilian amphipod (Orchestia Darwinii, fig. 8) presents a case of dimorphism, like that of Tanais; for there are two male forms, which differ in the structure of their chelæ. As either chela would certainly suffice to hold the female,—for both are now used for this purpose,—the two male forms probably originated by some having varied in one manner and some in another; both forms having derived certain special, but nearly equal advantages, from their differently shaped organs.

It is not known that male crustaceans fight together for the possession of the females, but it is probably the case; for with most animals when the male is larger than the female, he seems to owe his greater size to his ancestors having fought with other males during many generations. In most of the orders, especially in the highest or the Brachyura, the male is larger than the female; the parasitic genera, however, in which the sexes follow different habits of life, and most of the Entomostraca must be excepted. The chelæ of many crustaceans are