Page:Darwinism by Alfred Wallace 1889.djvu/142

 and recognition by members of the same variety or incipient species. It has also been observed that each differently coloured variety of wild animals, or of domesticated animals which have run wild, keep together, and refuse to pair with individuals of the other colours; and this must of itself act to keep the races separate as completely as physical isolation.

As natural selection acts solely by the preservation of useful variations, or those which are beneficial to the organism under the conditions to which it is exposed, the result must necessarily be that each species or group tends to become more and more improved in relation to its conditions. Hence we should expect that the larger groups in each class of animals and plants—those which have persisted and have been abundant throughout geological ages—would, almost necessarily, have arrived at a high degree of organisation, both physical and mental. Illustrations of this are to be seen everywhere. Among mammalia we have the carnivora, which from Eocene times have been becoming more and more specialised, till they have culminated in the cat and dog tribes, which have reached a degree of perfection both in structure and intelligence fully equal to that of any other animals. In another line of development, the herbivora have been specialised for living solely on vegetable food till they have culminated in the sheep, the cattle, the deer, and the antelopes. The horse tribe, commencing with an early four-toed ancestor in the Eocene age, has increased in size and in perfect adaptation of feet and teeth to a life on open plains, and has reached its highest perfection in the horse, the ass, and the zebra. In birds, also, we see an advance from the imperfect tooth-billed and reptile-tailed birds of the secondary epoch, to the wonderfully developed falcons, crows, and swallows of our time. So, the ferns, lycopods, conifers, and monocotyledons of the palæozoic and mesozoic rocks, have developed into the marvellous wealth of forms of the higher dicotyledons that now adorn the earth.

But this remarkable advance in the higher and larger groups does not imply any universal law of progress in organisation, because we have at the same time numerous examples (as has been already pointed out) of the persistence of lowly organised