Page:Darwinism by Alfred Wallace 1889.djvu/100

78 remarkable facts as to their variability. He declares that on the same branch of oak he has noted the following variations: (1) In the length of the petiole, as one to three; (2) in the form of the leaf, being either elliptical or obovoid; (3) in the margin being entire, or notched, or even pinnatifid; (4) in the extremity being acute or blunt; (5) in the base being sharp, blunt, or cordate; (6) in the surface being pubescent or smooth; (7) the perianth varies in depth and lobing; (8) the stamens vary in number, independently; (9) the anthers are mucronate or blunt; (10) the fruit stalks vary greatly in length, often as one to three; (11) the number of fruits varies; (12) the form of the base of the cup varies; (13) the scales of the cup vary in form; (14) the proportions of the acorns vary; (15) the times of the acorns ripening and falling vary.

Besides this, many species exhibit well-marked varieties which have been described and named, and these are most numerous in the best-known species. Our British oak (Quercus robur) has twenty-eight varieties; Quercus Lusitanica has eleven; Quercus calliprinos has ten; and Quercus coccifera eight.

A most remarkable case of variation in the parts of a common flower has been given by Dr. Hermann Müller. He examined two hundred flowers of Myosurus minimus, among which he found thirty-five different proportions of the sepals, petals, and anthers, the first varying from four to seven, the second from two to five, and the third from two to ten. Five sepals occurred in one hundred and eighty-nine out of the two hundred, but of these one hundred and five had three petals, forty-six had four petals, and twenty-six had five petals; but in each of these sets the anthers varied in number from three to eight, or from two to nine. We have here an example of the same amount of "independent variability" that, as we have seen, occurs in the various dimensions of birds and mammals; and it may be taken as an illustration of the kind and degree of variability that may be expected to occur among small and little specialised flowers.

In the common wind-flower (Anemone nemorosa) an almost equal amount of variation occurs; and I have myself gathered