Page:American Seashells (1954).djvu/46

28 males usually occupy positions in the vicinity of the female and move to the mating position at night. Occasionally bachelors are found which either by chance or choice remain solitary throughout the entire male phase.

Soon after hatching from the egg, and in one species (Crepidula adunca Sowerby from Panama) even before hatching, a slender copulatory organ, the verge or phallus, grows out from the body behind the right tentacle (fig. 7), As the female phase develops later in life, the verge begins to shrink and is finally absorbed as the female organs take form. Associated with these changes is a marked alteration in behavior, whereby the wandering individual, which was so characteristically masculine when young, now becomes strictly sedentary. She receives her mate, lays her eggs in capsules beneath her foot and broods her young until they are prepared for their own independence.

In our Common Slipper Shell, Crepidula fornicata, those individuals which live on muddy bottoms where there are no solid objects to which they can attach themselves, frequently pile up in groups of six to twelve or more. These groups continue from year to year, newly arrived young in the male phase attaching themselves to the top of the pile as the old, female-phase individuals die at the bottom.

Most marine prosobranchs, however, are of separate sexes (dioecious or unisexual). While some species in which the sex products of both sexes are discharged freely into the water have no outward morphological features, there are a great number of gastropods in which the male has an external copulatory organ or verge. The shape and position of the verge are often used in classifying families, genera or species.

Depending upon the species, and sometimes the genus, the females take care of their young in a variety of ways. In some there is no motherly instinct, and the eggs are liberated directly into the water where they float away on the chance of being fertilized by the free-swimming sperm from a nearby male. (See fig. 9 with Tectarius and Littorina.) In other types the eggs are fertilized and undergo development to the adult-like form in the uterine portion of the oviduct. Others have developed a kangeroo-like pouch in the tissues of their back where the young are allowed to develop to the adult form. Once liberated, however, the young do not return to the pouch. Viviparity or the giving birth to young alive (technically ovoviviparity) is known in Planaxis, Littorina saxatilis and a number of fresh-water species in several different families.

Among a large proportion of the marine gastropods, the females form special egg cases or capsules into which the eggs are placed, and where the